A gigantic explosion causes a massive tsunami to wash over Atlantis. The city's robotic sentinels erect an energy shield to protect the capital, but most citizens perish. The Atlantean Queen is drawn into a glowing blue beam and disappears, leaving her young daughter, Kida, behind. Atlantis sinks below the waves.
In 1914, Milo Thatch, an aspiring cartographer/linguist/explorer, works as a boiler operator in the basement of the Smithsonian Institution in Washington D.C.. Milo believes that his research has revealed the location of an ancient manuscript that allegedly reveals the location of Atlantis. Milo is greeted by Helga Sinclair, who invites Milo to see her employer. Her employer is Preston B. Whitmore, an eccentric millionaire who attended Georgetown University with Milo's grandfather, Thaddeus Thatch, in 1866. Preston wants Milo to translate the manuscript and lead an expedition to Atlantis. Milo agrees.
Milo sets out aboard the Ulysses, a submarine headed up by Commander Lyle Rourke, the military man, and Helga. Among the crew are Vincenzo 'Vinny' Santorini, the crew's Italian demolitions expert, Gaetan 'Mole' Moliere, a crazed French geologist, Dr. Joshua Sweet, the ship's medical officer, Audrey Ramirez, a teenaged tomboy mechanic, Jebidiah 'Cookie' Farnsworth, the ship's Western redneck cook, and Wilhelmina Packard, an elderly communications expert. As the team tries to find to a crevice that leads to an underground cavern leading to Atlantis, the Ulysses is attacked and destroyed by Leviathan. Milo, Rourke, and the remaining crew members escape using small sub-pods and three escape subs. They flee into the underground cavern they had been searching for. Along the way, all but one of the aqua evacs (carrying Sweet, Audrey, Cookie, Packard, Rourke, Helga, and several dozen crew) and one sub pod (carrying Mole and Vinny) are destroyed. The crew then proceeds to explore the cavern, unaware they are being tracked by a group of Atlanteans.
After their camp is destroyed by mysterious glowing insects, the team attempts to cross a bridge, but ends up at the bottom of a dormant volcano when the bridge collapses under the weight of the team's vehicles. Using his drilling machine (officially known as the Digger), Mole drills a tunnel out of the volcano. After doing so, the crew are met by Kida, now a young woman, albeit thousands of years old. Kida leads the group to her aging father, King Kashekim Nedakh, who tells them to leave. Rourke requests the King allow his team to stay the night in order to rest, and resupply, and the King grants it. Meanwhile, Kida enlists Milo's help in deciphering the runes throughout the city, which the Atlantean people can no longer read. She and Milo discover the nature of the Heart of Atlantis, but can not figure out how it works or where it is, because one of the pages is missing.
The Heart of Atlantis,
merged with Princess Kida
After some time, Milo and Kida are surrounded by Rourke along with his mercenaries; only to discover that Rourke reveals that he has the missing page, and that he and Helga intend to sell the Heart of Atlantis. Rourke then offers Milo to join him, so he can locate the crystal. Milo then challenges Rourke, stating that he isn't a mercenary; Rourke replies he'd rather be called an "adventure capitalist".
Rourke decides that he has had enough of Milo's resistance and instead proceeds to doing things his way; he forces Milo into being his translator and then confronts King Nedakh. When King Nedakh refuses to disclose the location of the Heart, Rourke fatally punches him. Rourke then forces Milo to find the Heart for him. Once found, the Heart reaches into Kida's mind. As Milo and Rourke argue, the Heart of Atlantis speaks through the possessed Kida, saying, "Soleshma-du note, Milo Thatchdu. Guan kene sus-ene", meaning "All will be well, Milo Thatch. Be not afraid." in Atlantean. Kida then strides toward the crystal, which levitates her up to it and merges with her, as Sinclair and Milo (Rourke is never surprised, as Vinnie had said) watch in surprise and are all equally shocked. Rourke locks Kida in a steel crate and prepares to leave for the caves.
When Rourke leaves Milo behind, Vinnie, Audrey, Mole, Cookie, and Packard, bothered by Rourke treating Milo and the Atlanteans harshly, decide to stay behind as well. King Nedakh, tells Milo about the Heart of Atlantis, explaining why the crystal has a mind of its own and the reason Atlantis sank beneath the ocean. The King then gives his crystal to Milo and tells him that his burden would have becomes Kida's when the time was right, but instead will fall to Milo. He tells Milo to save Atlantis and Kida, and dies.
Milo and the Atlanteans engage Rourke and his mercenaries inside the dormant volcano. A lengthy air battle ensues, and its the attacking Atlanteans with their flying "stone fish" against the defending mercenaries with their "Whitmore Wings". After the battle has been fought, Rourke and Helga are killed along with Rourke's mercenaries who either fled or died during the battle. However, the blast from Rourke's airship triggers a volcanic eruption that threatens to destroy the city. Milo returns to Atlantis with the Heart/Kida in tow. The Heart/Kida activates the city's sentinels, who erect an energy shield to stop the flow of lava. The Atlanteans reward the surviving crew and return them to the surface. The crew, now insanely wealthy, return to Whitmore's mansion in Washington and rehearse their cover story to hide the existence of Atlantis. Milo remains behind in Atlantis with Kida.
Voice cast
- Milo James Thatch: Michael J. Fox
- Kidagakash "Kida" Nedakh: Cree Summer
- Commander Lyle Tiberius Rourke: James Garner
- Gaetan "Mole" Moliére: Corey Burton
- Vincenzo "Vinny" Santorini: Don Novello
- Doctor Joshua Strongbear Sweet: Phil Morris
- Lieutenant Helga Katrina Sinclair: Claudia Christian
- Audrey Rocio Ramirez: Jacqueline Obradors
- King Kashekim Nedakh: Leonard Nimoy
- Preston B. Whitmore: John Mahoney
- Wilhelmina Bertha Packard: Florence Stanley
- Jedidiah Allardyce "Cookie" Farnsworth: Jim Varney
- Fenton Q. Harcourt: David Ogden Stiers
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